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1Evolutionarybiologistsbelievethatspeciation,theformationofanewspecies,oftenbeginswhensomekindofphysicalbarrierarisesanddividesapopulationofasinglespeciesintoseparatesubpopulations.Evolutionarybiologistsbelievethatspeciation,theformationofanewoftenbegins通常在….的时候开始whensomekindofphysicalbarrierarises当一些物理的出dividesapopulationofasinglespeciesintoseparatesubpopulations并且将一个All:不断发展的生物学家们相信这些新物种的形成通常在一些物理的出现并且将一个整2Physicalseparationbetweensubpopulationspromotestheformationofnewspeciesbecauseoncethemembersofonesubpopulationcannolongermatewithmembersofanothersubpopulation,theycannotexchangevariantgenesthatariseinoneofthesubpopulations.Physicalseparationbetweensubpopulationspromotestheformationofnewspecies在分组物种数量的科学分离会促进新物种的形成becauseoncethemembersofonesubpopulationcannolongermatewithmembersofanothersubpopulation因为当一个分组物种数量中的成员不可以再与另Theycannotexchangevariantgenes他们不可以交换不同的Thatariseinoneofthesubpopulations.3Wefindevidenceofsuchextendedeventsinthefossilrecord,whichaffordsglimpseintothebreakupofformerlycontinuousenvironments.Wefindevidenceofsuchextendedeventsinthefossilrecord我们在化石中找到了WhichaffordsglimpseintothebreakupofformerlycontinuousAll:我们在可以提供给我们看到之前环境的可能的化石中找到了那种扩张4Inothergroups,howevergeneticdivergenceshadnotproceededsofarandthedescendantscouldstillinterbreed–forthem,reproductiveisolationwasnotcompleted,andsospeciationhadnotoccurred.Inothergroups,however,geneticdivergenceshadnotproceededso不过,在其他的族群中,的并没有继续进andthedescendantscouldstillinterbreedForthem,reproductiveisolationwasnot对他们来说,生殖的并不彻Andsospeciationhadnotoccurred所以物种形成没有发生All:但是,在其他的族群中,的并没有继续进行,他们的后代还在互相繁殖,对于他们来说,生殖的并不彻底所以物种的形成没有发生。5AllopatricspeciationcanalsobebroughtbytheimperceptiblyslowbutcolossalmovementsofthetectonictesthatmakeupEarth’ssurface.AllopatricspeciationcanalsobebroughtbytheimperceptiblyslowA物种也可以被细微的slow带来ButcolossalmovementsofthetectonictesthatmakeupEarth’ssurface.All:A物种也可以被细微的slow6、GravesfoundthatallfourPacificenzymesfunctionbetteratlowertemperaturesthanthefourAtlanticversionsofthesameenzymes.All:Graves发现4个太平洋的酶在低温下的功能比四个一样的的酶更7.BecauseGraves’studyofgeographicallyisolatedpopulationsofisthmusfishesoffersaglimpseofthebeginningofaprocessofgradualaccumulationofmutationsthatareneutraloradaptive,divergencesheremightbeevidenceofallopatricspeciationinprocess.BecauseGraves’studyofgeographicallyisolatedpopulationsofisthmus因为gravesOffersaglimpseofthebeginningofaprocessofgradualaccumulationofmutations提供了突变的逐渐积累的过程的开始的一瞥thatareneutraloradaptive这是中立的、适合的Divergencesheremightbeevidenceofallopatricspeciationin这个在这里可能会成为过程中的分布区不All:因为grave8、Whilepreviouslythegapbetweenthecontinentshadallowedaflowofwater,nowtheisthmuspresentedabarrierthatdividedtheAtlanticOceanfromthePacificWhilepreviouslythegapbetweenthecontinentshadallowedaflowofNowtheisthmuspresentedabarrier现在这种海峡展现出了一种DividedtheAtlanticOceanfromthePacificOcean.这种将太平洋和分开All洋和分开的。9、Preschools–educationalprogramsforchildrenundertheageoffive–differsignificantlyfromonecountrytoanotheraccordingtotheviewsthatdifferentsocietiesholdregardingthepurposeofearlychildhoodeducation.Preschools–educationalprogramsforchildrenundertheageoffive–differsignificantlyfromonecountrytoanotheraccordingtotheviews学前班——一种为小于五岁的孩子提供的教育项目——在国家之间对此的看法有很大的差异Differentsocietiesholdregardingthepurposeofearlychildhoodeducation.不一样All10、Whereasparentsin tendtoseepreschoolsprimarilyasawayofgivingchildrenagoodstartacademically,Japaneseparentsviewthemprimarilyasawayofgivingchildrentheopportunitytobemembersofagroup.Whereasparentsin tendtoseepreschoolsprimarilyasawayofgivingchildrenagoodstartacademically中国的父母倾向与将学前班的作用看作给孩子良好的,Japaneseparentsviewthemprimarilyasawayofgivingchildrentheopportunitytobemembersofagroup.然而的父母把学前班的动机看作给孩子一个机会去体验团队,All:中国的父母倾向与将学前班的作用看作给孩子良好的学业基础,然而,的父母把学11、IntheUnitedStates,incomparison,parentsregardtheprimarypurposeofpreschoolsasmakingchildrenmoreindependentandself-reliantalthoughobtainingagoodacademicstartandhavinggroupexperiencearealsoimportant.12、Whilemanyprogramsdesignedforpreschoolersfocusprimarilyonsocialandemotionalfactors,somearegearedmainlytowardpromotingcognitivegainsandpreparingpreschoolersfortheformalinstructiontheywillexperiencewhentheystart13、WhetherHeadStartisseenassuccessfulornotdependsonthelensthroughwhichoneislooking.14Inaddition,resultsfromothertypesofpreschoolreadinessprogramsindicatethatthosewhoparticipateandgraduatearelesslikelytorepeatgrades,andtheyaremorelikelytocompleteschoolthanreadinessprogram,foreverydollarspentontheprogram,taxpayerssavedsevendollarsbythetimethegraduatesreachedtheageof15、Consequently,childrenofaparticularagecannotbeexpectedtomastereducationalmaterialwithouttakingintoaccounttheircurrentlevelofcognitive16、Thereisnosimpleorsinglefactorthatdeterminesifagivensitewillbeasavanna,butsomefactorsseemtoyimportantrolesintheirformation.17、Onetheorybehindsavannaformationisthatwet speciesareunabletowithstandthedryseason,andthussavanna,ratherthanrain,isfavoredonthe18Thoughplentyofrainmayfallonasavannaduringtheyearforatleastpartoftheyearlittledoes,creatingthedroughtstressultimayfavoringgrasses.19、SuchconditionsprevailthroughoutmuchofnorthernSouthAmericaandCuba,butmanyCentralAmericansavannasaswellascoastalareasofBrazilandtheislandofTrinidaddonotfitthispattern.20、Thoughmostsavannasarefoundonsiteswithpoorsoils(becauseofeithermoistureconditionsornutrientlevelsofboth),poorsoilscananddosupportlushrain.21、Frequentfireisafactortowhichrain speciesseemunabletoadapt,althoughancientcharcoalremainsfromAmazon soilsdatingpriortothearrivalofhumanssuggestthatmoistsalsooccasionallyburn.22、Whenburningoccurs,itpreventscompetitionamongntspeciesfromprogressingtothepointwheresomespeciesexcludeothers,reducingtheoveralldiversityoftheecosystem.23Oncethehumuslayerdisappearsnutrientscannotberecycledandleachfromthesoil,convertingsoilfromfertiletoinfertileandmakingitsuitableonlyforsavanna24、Frequentfireisafactortowhichrain speciesseemunabletoadapt,althoughancientcharcoalremainsfromAmazon soilsdatingpriortothearrivalofhumanssuggestthatmoistsalsooccasionallyburn.25、Evidencefromotherstudiessuggeststhatexclusionoffireresultsinmarkedlydecreasednt-speciesrichness,oftenwithanincreaseintreedensity.26、Successincolonizationdependstoagreatextentontherebeingasiteavailableforcolonization–asafesitewheredisturbancebyfireorbycuttingdownoftreeshaseitherremovedcompetingspeciesorreducedlevelsofcompetitionandothernegativeinctionstoalevelatwhichtheinvadingspecies e27、Afertile,plowedfieldisrapidlyinvadedbyalargevarietyofweeds,whereasaneighboringconstructionsitefromwhichthesoilhasbeencompactedorremovedtoexposeacoarse,infertileparentmaterialmayremainvirtuallyofvegetationformanymonthsorevenyearsdespitereceivingthesameinputofseedsastheplowed28Pioneerspeciesthosethatoccuronlyintheearlieststagesofcolonizationtendtohavehighratesofinvasionbecausetheyproduceverylargenumbersofreproductivepropagules(seeds,spores,andsoon)andbecausetheyhaveanefficientmeansofdispersal(normally,wind)29、Manyntsdependonwindfordispersalandproduceabundanttiesofsmallrelativelyshort-livedseedstocompensateforthefactthatwindisnotalwaysareliablemeansofreachingtheappropriatetypeofhabitat.30Thisisparticularlyimportantforspeciesthatcolonizeanenvironmentwherethereisnoexistingvegetationtoameliorateclimaticextremesandinwhichtheremaybegreatclimaticdiversity.31Thefirstcommunitythatoccupiesadisturbedareathereforemaybecomposedofspecieswiththehighestrateofinvasion,whereasthecommunityofthesubsequentstagemayconsistofntswithsimilarsurvivalratesbutlowerinvasionrates.32Thesefamilieswerepensionedoffgivenasumofmoneytoliveon)orsimplyhadtheirrevenuestakenawayorrestricted;theirsonswereenticedawaytoschoolsfordistrictofficers,latertobepostedinsomefarawayprovince;andtheoldpatron-relationsthathadboundtogetherlocalsocietiessimplydisintegrated.33、TheaverageannualvolumeofriceexportedfromSiamgrewfromunder60millionkilogramsperyearinthelate1850stomorethan660millionkilogramsperyearattheturnofthecentury;andoverthesameperiodtheaveragepriceperkilogramdoubled.34、Thisgrowthwasachieveastheresultofthecollectivedecisionsofthousandsofpeasantsfamiliestoexpandtheamountoflandtheycultivated,clearandntnewland,oradoptmoreintensivemethodsofagriculture.35Undertheseconditions,itmadegoodsensetothousandsofpeasantfamiliestoineffectworkfull-timeatwhattheyhadbeenabletodoonlypart-timepreviouslybecauseoftherequirementtoworkfortheernment:growriceforthemarketce.36Bytheearlytwentiethcenturyasernmentmodernizationinasensecaughtupwithwhathadbeenhappeninginthecountrysidesincethe1870s,theernmentbureaucracyintrudedmoreandmoreintovillagelife.37、HubbellandJohnsonobservedthatsomespeciesofstinglessbeesarehighlyaggressivetomembersoftheirspeciesfromothercolonies,whileotherspeciesare38、ThoughHubbellandJohnsonwereinterestedinhowbeebehaviormightaffectcolonydistributions,theyrecognizedthattheavailabilityofpotentialnestsitesforcoloniescouldalsoaffectdistributions.39Thenumberofcoloniesinthestudyareawasnotlimitedbyavailabilityofsuitabletrees,andaclumpedorregulardistributionofcolonieswasnotduetoanunderlyingclumpedorregulardistributionofpotentialnestsites.40、Whilethiscontestoveranunoccupiednestsiteproducednoobviousmortality,fightsoveroccupiednestssometimeskillover1,000beesinasinglebattle.41、Manyprehistoriansbelievethatfarmingmayhaveemergedindependentlyinseveraldifferentareasoftheworldwhensmallcommunities,drivenbyincreasingpopulationandadeclineinavailablefoodresources,begantontseedsinthegroundinanefforttoguaranteetheirsurvival.42、Anadditionalbenefitwasthatthesedimentcarriedbytheriverwatersdepositednutrientsinthesoil,thusenablingthefarmertocultivateasingleplotofgroundformanyyearswithoutmovingtoanewlocation.43、Astimewenton,suchcommunitiesgraduallylearnedhowtodirecttheflowwatertoenhancetheproductivecapacityoftheland,whiletheintroductionoftheironploweventuallyledtothecultivationofheavysoilsnotpreviouslysusceptibleto44、Theincreaseinfoodproductionintheseregionsledtoasignificantgrowthinpopulation,whileeffortstocontroltheflowofwaterto izetheirrigationofcultivatedareasandtoprotectthelocalinhabitantsfromhostilesoutsidethecommunityprovokedthefirststepstowardcooperativeactivitiesonalargescale.45、Whilesomeoftheseurbancenterswereidentifiedwithaparticulareconomicfunction,suchasproximitytogoldorirondepositsorastrategiclocationonamajortraderoute,othersservedprimarilyasadministrativecentersorthesiteoftemplesfortheofficialcultorotherritualobservances.46、Insomecases,thephysicaldivisionwithinthefirstcitiesreflectedthestricthierarchicalcharacterofthesocietyasawhole,witharoyalpalacesurroundedbyanimposingwallandseparatefromtheremainderoftheurbanpopulation.47、NativeAmericanshadgrowncornintheregionforyearsbutneverinsuchlargefieldsasdidlatersettlerswhobecamefarmers,whosesurpluseswereshippedeast.48、Intheabsenceofthesefires,treesreappearedonlandnotincultivationand,undisturbed,eventuallyformed49、Theearlierunbrokenlandscapegavewaytoindependentfarms,eachfencedoffinaprecisecheckerboardpattern.50、Howevernogrouporspeciescanmaintainitsdominanceindefiniyandwhen,afterover200millionyears,theageofdinosaurscametoadramaticendabout65millionyearsago,mlsbegantoflourish,evolvingfromrelativelyfewtypesofsmallterrestrialanimalsintothemyriadofdiversespecies,includingbatsandwhales,thatweknowtoday.51、Anextremelyabundantsetofnktonicmarineanimalscalledforaminiferalargelydisappeared,althoughtheyreboundedlater.52、Formanyyearsscientistsassumedthatacoolingoftheclimatewasresponsible,withdinosaursbeingparticularlyvulnerablebecause,likemodernreptiles,theywereectothermic(dependentonenvironmentalheat,orcold-blooded).53Nevertheless,climaticexnationsfortheK-Textinctionarenotreallychallengedbytheideasthatdinosaursmayhavebeenendothermic,becauseevenendothermscanbeaffectedbyasignificantchangeintheclimate.54、ExnationsfortheK-Textinctionwererevolutionizedin1980whenagroupofphysicalscientistsledbyLuisAlvarezproposedthat65millionyearsagoEarthwasstuckbya10-kilometer-widemeteoritetravelingat90,000kilometersperhour.55、ThereforeAlvarezandhiscolleaguesconcludedthatitwaslikelythattheiridiuminsedimentaryrocksdepositedattheK-Tboundaryhadoriginatedinagiantmeteoriteorasteroid.56、Mostscientistcametoacceptthemeteoritetheoryafterevidencecametolightthatacircularformation,180kilometersindiameterandcenteredonthenorthcoastoftheYucatanPeninsula,wascreatedbyameteoriteimpactabout65millionyears57、Thiscameaboutfromtwomajordevelopments:paperbecamecommon,recingparentasthemajormediumforwriting,androundedscriptswereregularizedandperfectedsothattheyrecedtheangularscriptsofthepreviousperiod,whichbecauseoftheirangularitywereuneveninheight. toinclude,ofcourse,manuscriptsoftheKoran,whicheveryMuslimwantedtoread,butscientificworks,histories,romances,andepicandlyricpoetrywerealsocopiedinfinehandwritinganddecoratedwithbeautifulillustrations.59、Papermakinghadbeenintroducedtotheclandsfrom intheeighthcentury.Ithasbeensaidthat papermakerswereamongtheprisonerscapturedinabattlefoughtnearSamarqandbetweenthe andtheMuslimsin751,andthetechniqueofpapermaking–inwhichcellulosepulpextractedfromanyofseveralntsisfirstsuspendedinwater,caughtonafinescreen,andthendriedintoflexiblesheets–slowlyspreadwestward.60、PapermakingspreadquicklytoEgyptandeventuallytoSicilyandSpainbutitwasseveralcenturiesbeforepapersupntedparentforcopiesoftheKoran,probablybecauseoftheconservativenatureofreligiousartanditspractitioners.61、Althoughpaperwasneverascheapasithas etoday,itwasfarlessexpensivethanparent,andthereforemorepeoplecouldaffordtobuybooks,Paperisthinnerthanparent,somorepagescouldbeenclosedwithinasingle62、Atfirstpaperwasmadeinrelativelysmallsheetsthatwerepastedtogetherbutbythebeginningofthefourteenthcentury,verylargesheets–asmuchasameteracross–wereavailable.63、Thisinturnallowedtheeasytransferoficideasandmotifsoverdistancesfromonemediumtoanother,andinadifferentscaleinwaysthathadbeendifficult,ifnotimpossible,inthepreviousperiod.64、RoundedstylesofArabichandwritinghadlongbeenusedforcorrespondence salongsidetheformalangularscriptsusedforinscriptionsandmanuscriptsoftheKoran.65、ButRussia’spotentialforgrowthwasli
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